Tan, Ge, and Boris Lenhard. 2016. “TFBSTools: An R/Bioconductor Package for Transcription Factor Binding Site Analysis.” Bioinformatics , January. Oxford Univ Press. doi:10.1093/bioinformatics/btw024.

Harmston, Nathan, Elizabeth Ing-Simmons, Ge Tan, Malcolm Perry, Matthias Merkenschlager, and Boris Lenhard. 2016. “Topologically Associated Domains Are Ancient Features That Coincide with Metazoan Clusters of Extreme Noncoding Conservation.” bioRxiv. doi:10.1101/042952.


Harmston, Nathan, Elizabeth Ing-Simmons, Malcolm Perry, Anja Barešić, and Boris Lenhard. 2015. “GenomicInteractions: An R/Bioconductor Package for Manipulating and Investigating Chromatin Interaction Data.” BMC Genomics 16 (1). biomedcentral.com: 963.

Haberle, Vanja, Alistair R. R. Forrest, Yoshihide Hayashizaki, Piero Carninci, and Boris Lenhard. 2015. “CAGEr: Precise TSS Data Retrieval and High-Resolution Promoterome Mining for Integrative Analyses.” Nucleic Acids Research 43 (8). Oxford Univ Press: e51.

Ing-Simmons, Elizabeth, Vlad C. Seitan, Andre J. Faure, Paul Flicek, Thomas Carroll, Job Dekker, Amanda G. Fisher, Boris Lenhard, and Matthias Merkenschlager. 2015. “Spatial Enhancer Clustering and Regulation of Enhancer-Proximal Genes by Cohesin.” Genome Research 25 (4): 504–13.


FANTOM Consortium and the RIKEN PMI and CLST (DGT), Alistair R. R. Forrest, Hideya Kawaji, Michael Rehli, J. Kenneth Baillie, Michiel J. L. de Hoon, Vanja Haberle, et al. 2014. “A Promoter-Level Mammalian Expression Atlas.” Nature 507 (7493): 462–70.

Haberle, Vanja, Nan Li, Yavor Hadzhiev, Charles Plessy, Christopher Previti, Chirag Nepal, Jochen Gehrig, et al. 2014. “Two Independent Transcription Initiation Codes Overlap on Vertebrate Core Promoters.” Nature 507 (7492). Nature Publishing Group: 381–85.

Zuin, Jessica, Vedran Franke, Wilfred F. J. van Ijcken, Antoine van der Sloot, Ian D. Krantz, Michael I. J. A. van der Reijden, Ryuichiro Nakato, Boris Lenhard, and Kerstin S. Wendt. 2014. “A Cohesin-Independent Role for NIPBL at Promoters Provides Insights in CdLS.” PLoS Genetics 10 (2): e1004153.


Harmston, Nathan, Anja Baresic, and Boris Lenhard. 2013. “The Mystery of Extreme Non-Coding Conservation.” Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences 368 (1632): 20130021.

Harmston, Nathan, and Boris Lenhard. 2013. “Chromatin and Epigenetic Features of Long-Range Gene Regulation.” Nucleic Acids Research 41 (15): 7185–99.


Lenhard, Boris, Albin Sandelin, and Piero Carninci. 2012. “Metazoan Promoters: Emerging Characteristics and Insights into Transcriptional Regulation.” Nature Reviews. Genetics 13 (4): 233–45.

Ragvin, Anja, Enrico Moro, David Fredman, Pavla Navratilova, Øyvind Drivenes, Pär G. Engström, M. Eva Alonso, et al. 2010. “Long-Range Gene Regulation Links Genomic Type 2 Diabetes and Obesity Risk Regions to HHEX, SOX4, and IRX3.” Proceedings of the National Academy of Sciences of the United States of America 107 (2): 775–80.

Akalin, Altuna, David Fredman, Erik Arner, Xianjun Dong, Jan Christian Bryne, Harukazu Suzuki, Carsten O. Daub, Yoshihide Hayashizaki, and Boris Lenhard. 2009. “Transcriptional Features of Genomic Regulatory Blocks.” Genome Biology 10 (4): R38.

A complete list of publications is available on our Imperial College page.